A Listing of Fossil Sharks and Rays of the World

Copyright 1999 George S. Williams

Introduction

Euselachii

Neoselachii

Batomorphii

References

Batomorphii CAPPETTA, 1980 (Skates and Rays)

Rajiformes BERG, 1940

Uncertain Family

Erguitaia arganiae (ARAMBOURG, 1952) ‑ Maastr.: Oued Erguita, Morocco

benningensis CASE et al, 2001 – Sant.: Chattahoochee Co., GA

rugos CASE et al, 2001 – Sant.: Chattahoochee Co., GA

sp. – Maastr.: Charquieh, Syria

Ewingia problematica “decurrens” CASE & CAPPETTA, 1997 ‑ Maastr.: Hunt Co., TX

Germanobatos ornatus MUELLER & DIEDRICH, 1991 - Cen.: Westfalen, Germany

Tanoutia iminensis CAPPETTA & NOUBHANI, 1995 - Maastr.: Morocco

Turoniabatis cappettai LANDEMAINE, 1999‑ Cen.: Lussant, France

Archaeobatidae DELSATE & CANDONI, 2001

Cristabatis crescentiformis DELSATE & CANDONI, 2001 – Mid.Toa.: France

exundans DELSATE & CANDONI, 2001 – Up.Toa.: Nevers, France

Doliobatis weisi DELSATE & CANDONI, 2001 – Up.Toa.: France

Toarcibatis alticarinata DELSATE & CANDONI, 2001 – Up.Toa.: Nevers, France

brevicastata DELSATE & CANDONI, 2001 – Low.Toa.: France

elongata DELSATE & CANDONI, 2001 – Mid.Toa.: France

multicristata DELSATE & CANDONI, 2001 – Low.Toa.: France

sp. – Sin.: AZ

Cyclobatidae CAPPETTA, 1980

Cyclobatis longicanudatus - Up.Cret.: Lebanon

major DAVIS, 1887 - Cen.: Hakel/Hadjula, Lebanon

oligodactylus EGERTON, 1844 - Cen.: Hakel, Lebanon

tuberculatus CAPPETTA, 1980 - Cen.: Hakel, Lebanon

Gymnuridae FOWLER, 1934 (Butterfly Rays)

Gymnura delpiti CAPPETTA, 1984 ‑ Tha.: Khouribga, Morocco

grootaerti HERMAN, 1984 – Tha./Ypr.: Dormaal, Belg.; Ypr.: Egem, Belg.

hovestadti HERMAN, 1984 - Olig.: Tongeren, Belg.

laterialata WERNER, 1989 - Up.Cen.: Bahariya, Egypt

ssp. – Pal.: Morocco; Mid:Eoc.: Bracklesham, England; Togo; Mio.: Bakersfield, CA; Malta; Loupian, France; BajaCalifornia,

Mexico

Jacquhermania duponti (WINKLER, 1874)‑ Ypr.: Egem, Belg.; Meridian, MS; Stafford Co., VA; Mid:Eoc.: Bracklesham, England;

Lut.: Vald’Oise, France

Ouledia sigei CAPPETTA, 1986- Ypr.: Morocco

sp. – Tha../Ypr.: Senegal; Ypr.: Mali

Hypsobatidae CAPPETTA, 1992

Angolaia benguelaensis ANTUNES & CAPPETTA, 2002 – Up.Camp../Maastr.: Angola

Baibishia babishe NESOV & UDOVITSHENKO, 1986 - Low.San.: Kansay, Tajikistan

Hypsobatis weileri CAPPETTA, 1992 - Maastr.: Arad, Israel; Egypt; Morocco

sp. - Camp.: Negev Desert, Israel; Maastr.: Syria

Youssoubatis ganntourensis CAPPETTA, 1992 ‑ Maastr.: Khouribga, Morocco

sp. ‑ Maastr.: Arad, Israel; Syria

Parapaleobatidae CAPPETTA, 1992

Parapaleobates atlanticus ARAMBOURG, 1952 – Up.San./Maastr.: Jordan; Camp.: Morocco; Maastr.: Tanout, Morocco;

Charquieh, Syria

pygmaeus (ZITTEL, 1902) - San.: Wadi-Um-Ghudar, Jordan; France; Maastr.: Dachel, Egypt; Algeria; Iraq; Zaire

sp. – Up.Maastr.: Berivotra, NW Madagascar

Platyrhinidae JORDAN, 1923

Myledaphus bipartitus COPE,1876 ‑ Tur.: Grant Co., SD; Camp.: Blaine Co., MT; Big Horn River, WY; Camp./Maastr.: Milk River,

Alberta, Canada; Maastr.: Weld Co., CO; Garfield Co., MT; Slope Co., ND; Carbon/Niobrara Co., WY; NM; ND;

Pal.: McCone Co., MT

ssp. – Low.Cret.: CO; Camp.: Wheatley Co., MT: Kane Co., UT; Mayberries, Alberta, Canada; Up.Cret.: Holmdel, NJ;

Low. Pal.: Huxley/Alberta, Canada; Low.Eoc.: CO; WY

tritus NESSOV, 1981- Up.Cret.: Kizylkhum Desert, Uzbekistan

Platyrhina bolcensis MOLIN, 1860 - Up.Cret.: Monte Bolca, Italy

dockeryi CASE, 1994 – Ypr.: Meridian, MS

egertoni (ZIGNO, 1876) - Up.Cret.: Monte Bolca, Italy

sp. – Camp./Maastr.: Nardo, Italy

Pseudohypolophus ellipsis CASE et al, 2001 – Sant.: Chattahoochee Co., GA

mcnultyi THURMOND, 1971‑ Hau.: Unienville, France; Low.Brm.: Teruel, Spain; Cen.:Lussant, France;

Apt.: Hood Co., TX; Cen./Tur.: Navajo Co., AZ; Tur.: Grayson Co., TX; Con.: Tarrant Co., TX; Con./San.: Bernalillo

Co., NM; San.: Wadi-Um-Ghudar, Jordan; Camp.: Newcastle Co.,DE ; Camp./Maastr.: Hale/Montgomery Co., AL;

Monmouth Co., NJ; Maastr.: Bowie, MD: Wayne Co., NC; ND

Potamotrygonidae GARMAN, 1877 [Fresh Water Stingrays]

Potamotrygon ssp - Up.Mio.: Parana Rvr., Entre Rios Prov., Argentina

Pristidae BONAPARTE, 1838 [Sawfish]

Anoxypristis “Pristis” ensidens LEIDY, 1877- Neogene: NJ

fajumensis (STROEMER, 1903) - Mid./Up.Eoc.: Birket-el-Qurong, Egypt; Olig.: Summerville, SC

ferrinus BOHM, 1926 - Eoc.: Africa

imhoffi LERICHE, 1932 - Lut.: Vald’Oise, France; Belg.

malembeensis DARTEVILLE & CASIER, 1943 - Mio.: Zaire

mucrodens WHITE, 1926 ‑ Ypr.: Stafford Co., Va; Eoc.: Khouribga, Morocco; Nigeria; Mid./Up.Eoc.: Birket-el-Qurong, Egypt

priemi LERICHE, 1932 - Eoc.: France

ssp. - Ypr.: Stafford Co., VA; Mid:Eoc.: Bracklesham, England; Lut.: Kizylkum Desert, Uzbekistan

Peyeria libyca WEILER, 1935 - Cenom..: Baharija, Egypt; Gara es Sabaa, Morocco

Pristis agassizi GIBBS, 1847 - Eoc.: SC

amblodon COPE, 1869 - Mid.Eoc.: NJ

aquitanicus DELFORTREI, 1872 ‑ Mio.: Loupian, France

atlanticus ZBYSZENINSKY, 1947 - Mid. Mio.: Portugal

brevis CASIER, 1949 - Eoc.: Belg.

caheni DARTEVILLE & CASIER, 1959- Mio.: W.Africa

hamatus WHITE, 1926 – Tha.etian/Ypr.: Morocco

lathami “brayi” GALEOTTI, 1837 ‑ Ypr.: Covington Co., AL; Stafford Co. VA; Morocco; Lut.: Balegem, Belg.; Guitry, France;

Eoc.: Duplin Co, NC; Mid:Eoc.: Bracklesham, England; Up.Eoc.: Twiggs Co., GA; Togo;

Mid./Up.Eoc.: Birket-el-Qurong, Egypt

olbrechtsi DARTEVILLE & CASIER, 1959 - Mid.Eoc.: W. Africa

pectinatus (LATHAM, 1794) - Mio.: FL; Plio.: Beaufort Co., NC

pickeringi CASE, 1981- Up.Eoc.: Twiggs Co., GA

praecursor CASIER, 1946 - Eoc.: Belg.

propinquidens CASIER, 1949 - Eoc.: Belg.; Neth.

prosulcatus STROEMER, 1905 - Eoc.: Egypt

ssp. – Tha.: Meridian, MS; Lut.: Kizylkum Desert, Uzbekistan; Lut./Plio.: BajaCalifornia, Mexico; Eoc.: Rose Hill, NC;

Monroe Co., AL; Olig./Mio.: NC; Low.Mio.: NW Venezuela; Mio.: Sarasota Co., FL; Switzerland;

Plio.:Beaufort Co., NC; Pleist.: Hillsborough Co., FL; Java

Propristis schweinfurthi DAMES, 1883 ‑ Ypr.: Stafford Co., VA; Eoc.: Gordon, GA; LA; Cabinda, Nigeria; Togo; Mid:Eoc.: Bracklesham, England; Birket-el-Qurong, Egypt; Olig.: Clarke Co., AL; BajaCalifornia, Mexico

ssp. - Tha.: France; Tha../Ypr.: Meridian, MS; Ypr.: Morocco

Rajidae BONAPARTE, 1831 [Skates]

Mafcletia tibiensis WERNER, 1989 - Up.Cen.: Bahariya, Egypt

Raja antiqua LERICHE, 1926 - Plio.: Kallo, Belg.

binoculata (GIRARD, 1854) – Mio.: Santa Cruz Co., CA; Plio.: Hunboldt/Curry Co., OR;

casieri STEURBAUT & HERMAN, 1978 - Olig.: Antwerp, Belg.; Espenheim, Germany

cecilae “heinzelini” “terhagensis” STEURBAUT & HERMAN, 1978- Olig.: Espenhein, Germany; Szczein, Poland; Belg.

clavata LINNAEUS, 1758 ‑ Plio.: Kallo, Belg.

? davisi FOWLER, 1958 – Up.San.: Sahel Alma, Lebanon

“Pararaja” “Rajorhina” expansa (DAVIS, 1887) - Cen.: Hakel, Lebanon

farishi CASE & CAPPETTA, 1997 ‑ Maastr.: Hunt Co., TX

gentili (JOLEAUD, 1912) – Lan.: Bonpas, France; Srv.: Malta; Mio.: Switzerland

harrisae WARD, 1984 - Ypr.: Essex, England; Wales

kincaidi (GARMAN, 1908) – Plio.: Curry Co., OR

louisi CAPPETTA, 1972 - Up.Pal.: Nigeria

olisiponensis (JONET, 1968) ‑ Srv.: Cabrieres, France; Tort.: Caparica, Portugal; Up.Mio.: Liessel, Neth.

praealba ARAMBOURG, 1952 - Ypr.: Ouled Abdoun, Morocco

ssp. – Camp.: Aachen, Germany; Tha.: Berkeley Co., SC; MS; Ypr.: Sheppey, England; Up.Eoc./Plio.: BajaCalifornia, Mexico;

Mid.Eoc.: Seymour Island; Antarctica; Eoc.:Helmstedt, Germany; Up.Eoc./Olig.: OR; Olig.: Summerville, SC;

Espenhein, Germany; Szczein, Poland;Czeck.; Low.Mio.: Kent Co., DL; Tortonian: Portugal; Up.Mio.: MD; N.Germany;

Mio.: FL; Ishikawa, Japan; France; Switzerland; Mio./Plio.: Cape Province, South Africa;

Low.Plio.: Cubagua, Venezuela; Plio.: Beaufort Co., NC; Belg.; S.France; Italy; Pleist.: Orange Co., CA

sudhakari PRASSAD & CAPPETTA, 1993- Maastr.: Asifabad, India

Sechmetia "Squatina" aegyptica (STROEMER, 1927) - Up.Cret.: Egypt

cruciformes WERNER, 1989 - Up.Cen.: Bahariya, Egypt

Smithraja forestensis HERMAN, 1986 - Ypr.: Egem, Belg.

Walteraja exigua SIVERSON & CAPPETTA, 2001 – Low.Maastr.: Balsvik Quarry, Sweden

Rhinobatidae MUELLER & HENLE, 1838

Arthropterus rileyi AGASSIZ, 1843 - Lias.: Lyme Regis, England

Ataktobatis variabilis CAPPETTA & CORRAL, 1999 – Maastr.: Egypt; Morocco; Cantabria, Spain; Charquieh, Syria

Belemnobatis kemacki UNDERWOOD & WARD, 2004 – Bth.: SW England

morinicus (SAUVAGE, 1873) - Portlandian: Pas-de-Calais, France; Up.Jur.: Germany

sismondae THOLLIERE, 1854 - Kim.: Cerin, France; Tithonian: Nusplingen, Germany

sp. – Aal.: France; Mid.Bth.: SW England

stahli UNDERWOOD & WARD, 2004 – Bth.: SW England

variabilis REES, 2002 - Ber.: Pubeck, Southern England

"Spathobatis" werneri (THIES, 1983) – Kim.: Normandy, France; Tithonian: Charente-Maritime, France

Breviacanthus brevis (PHILLIPS, 1871) - Bath.: Stonesfield, England; Up.Jur.: Germany

Engaibatis schultzei ARRATIS, KRIWET & HEINREICK, 2002 - Up.Jur.: Tendaguru, Tanzania

Euryarthra moenstri AGASSIZ, 1843 - Tithonian: Solnhofen, Germany

Hamrabatis ornata - Maastr.: Egypt

sp. – Maastr.: Charquieh, Syria

weltoni CASE & CAPPETTA, 1997 - Maastr.: Hunt Co., TX

Iansan beurlini (SANTOS, 1968) - Apt.: NE Brazil

Isidobatus tricarinatus WERNER, 1989 - Up.Cen.: Bahariya, Egypt

Microbatis sabatieri CAPPETTA, 1983 – Tha.: Ouarzazate, Morocco

Platyrhinoides triseriata (JORDAN & GILBERT, 1880) – Pleist.: Orange Co., CA

Protoplatyrhina hopii WILLIAMSON, KIRKLAND & LUCAS, 1993 - Up.Cen./Mid.Tur.: Black Mesa, Navajo Co., AZ

renae CASE, 1978 ‑ Tur.: Grant Co., SD; San.: Chattahoochee Co., GA; Bernalillo Co., NM; Camp.: Sandoval Co., NM;

Worland, WY; Blaine Co.,MT; Alberta, Canada; Chihuahua, Mexico; Maastr.: Monmouth Co., NJ; Hunt Co., TX

Rhinobatos antunesi (JONET, 1968) ‑ Mio.: Loupian, France; Portugal

berytensis SIGNEAUX, 1951 ‑ Maastr.: Khouribga, Morocco

bruxelliensis (JAEKEL, 1894) – Tha../Ypr.: Merician, MS; Dormaal, Belg.; Morocco; Ypr.: Stafford Co., VA; Egem, Belg.;

Mid.Eoc.: Pamunkey River, VA; Bracklesham, England; Lut.: Guitry, France

casieri HERMAN, 1977- San.: Lonzee, Belg.; Wadi-Um-Ghudar, Jordan; Camp.: Newcastle Co.,DE ; Hunt Co., TX; Blaine Co., MT;

Milk River, Alberta, Canada; Up.Camp.: Washakie Co., WY; Up.Maastr.: Monmouth Co., NJ; Up.Eoc.: Twiggs Co., GA

craddocki CASE & CAPPETTA, 1997 ‑ Maastr.: Hunt Co., TX

echevei CAPPETTA & CORRAL, 1999 - Maastr.: Condado de Trevino, Cantabria, Spain

grandis (DAVIS, 1887) - Cen.: Hakel, Lebanon

hakelensis CAPPETTA, 1980 - Up.Cret.: Lebanon

halteri BIDDLE & LANDEMAINE, 1989 – Brm.: Unienville, France; Alb.: St. Dizier, France

ibericus CAPPETTA & CORRAL, 1999 - Maastr.: Condado de Trevino, Cantabria, Spain

incedens KRIWET, 1999A - Low.Brm.: Alcaine/Teruel, Spain

incertus CAPPETTA, 1973 ‑ Cen.: Baca Co., CO; Russel Co., KS; Fall River Co., SD; Tur.: Butte Co., SD;

Maastr.: Hunt Co., TX

intermedius - Up.Cret.: Lebanon

kiestensis CASE & CAPPETTA, 1999 - Tur./Cen.: Dallas Co., TX

landoniaensis CASE & CAPPETTA, 1999 - Camp.: Fanin/Lamar Co., TX

latus (DAVIS, 1887) - San.: Sahel Alma, Lebanon

lobatus CASE & CAPPETTA, 1999 - Tur./Cen.: Dallas Co., TX

maronita (PICTET & HUMBERT, 1866) - Cen.: Switzerland; Hakel, Lebanon

matzensis BAULT & GENAULT, 1995 - Tha.: Denmark, Ressons sur Matte, France; Tha./Ypr.: Dormaal, Belgium

obtusus (COSTA, 1864) - Apt.: Naples, Italy

palaeoceous ZHELSKO, 1999 - Up. Cret.: Crimea, Mangyshlak

picteti CAPPETTA, 1975 - Low.Brm.: Teruel, Spain; Apt.: Vaucluse, France

primarmatus (WOODWARD, 1889) - San.: Sahel Alma, Lebanon

productus GIRARD, 1855 – Pleist.: Orange Co., CA

ssp. – Toa.: Nevers, France; Kim.: Shewa, Africa; Alb.: Wissant, France; Tur.: Socorro Co., NM; Cen./Tur.: Navajo Co., AZ;

Con.: Saskatchewn, Canada; Camp.: Sandoval Co., NM; Hemmoor/Westfalen, Germany; Maastr.: ND; Benguerir,

Morocco; Lleida, Spain; Neth.; Syria; Nafud, Saudia Arabia; Tha.: Oise, France; Up.Pal.: Meridian, MS.; Ingall, Niger;

Ypr.: Stafford Co., VA; Egem, Belg.; Solan Dist., India; Morocco; Togo; Lut.: Covington Co., AL; Mid.Eoc.: Gerecse,

Hungary; Bart/Plio..: BajaCalifornia, Mexico; Eoc./Plio.: San Diego Co., CA; Haut.: Unienville, France;

Mid.Mio.: Beaufort Co., NC; Mio.: Kern Co., CA; Pelican Point, FL; Liessel, Neth

steurbauti CAPPETTA & NOLF, 1981- Lut.: Vald’Oise, France; Kizylkum Dsrt., Uzbekistan; Bart.: Ronquerolles, France

tesselatus VON DER MARCK, 1894 - Camp.: Westfalen, Germany

tenuirostris (DAVIS, 1887) - San.: Sahel Alma, Lebanon

uvulatus CASE & CAPPETTA, 1997 ‑ Con.: Denton Co., TX; Camp.: Fanin Co., TX; Maastr.: Hunt Co., TX

whitfieldi (HAY) - Cen.: Hadjula, Lebanon

Rhombopterygia rajoides CAPPETTA, 1980 - Camp.: Westfalen, Germany; Cen.: Hadjula, Lebanon

Spathobatis bugesiacus "Asterodermus platypterus" THOLLIERRE, 1854 - Kim.: Dorsett, England; Cerin, France;

Tithonian: Solnhofen Germany

desaltei UNDERWOOD & WARD, 2004 – Bth.: SW England

halteri BIDDLE & LANDEMAINE, 1989‑ Low.Brm.: Teruel, Spain; Alb.: Kent, England

moorbergensis SAUVAGE, 1873 – Toa.: Nevers, France; Jurrasic: Moorberg, Germany; Luxemborg

mutterllosei THIES, 1983 – Kim.: Normandy, France; Tithonian: Charente-Maritime, France

rugosus UNDERWOOD, MITCHELL & VELTKAMP, 1999 - Hau.: Speeton, Yorkshire, England

sp. – Brm.: Unienville, France; Kim.: Normandy, France; Tithonian: Charente-Maritime, France

uppensis THIES, 1983 - Kim.: Normandy, France; Hanover, N.Germany

Vascobatis albaifensis CAPPETTA & CORRAL, 1999 - Maastr.: Condado de Trevino, Cantabria, Spain

Rhynchobatidae GARMAN, 1913

Rhynchobatus pristinus (PROBST, 1877) ‑ Mio.: Craven Co., NC; Loupian, France; Germany; Malta; Portugal

ssp. - Maastr.: Morocco; Pal.: Ingall, Niger; Ypr.: Senegal; Mid.Eoc.: Pamunkey River, VA; Eoc.: Togo;

Olig.: Summerville SC; Mio.: Alachua Co., FL; Beaufort Co., NC; Mizunami, Japan; Malta; Mio./Low.Plio.: Venezuela; Pleist.: Hillsborough Co., FL

vincenti JAEKEL, 1894 – Eoc.: Lee‑on‑the‑Solent, England; Guineau-Bissau; Togo;Ypr.: Egem, Belg.; Lut.: Vald’Oise, France

Sclerorhynchidae CAPPETTA, 1974 [Sawfish]

Ankistrorhynchus lonzeensis CASIER, 1964 ‑ San.: Lonzee, Belg.

major CASE & CAPPETTA, 1999 - Camp.: Hop Brook, NJ

washakiensis CASE, 1987 - Up.Camp.: Washakie Co., WY

Baharipristis bastetiae WERNER, 1989 - Up.Cen.: Bahriya, Egypt

Borodinopristis ackermani CASE et al, 2001 - San.: Chattahoochee Co., GA

schwimmeri CASE, 1987 ‑ Camp.: Stewart Co.,GA

sp. ‑ Eoc.: Gordon Co., GA

Celtipristis herreroi KRIWET, 1999A Low.Brm.: Alcaine/Teruel, Spain

Ctenopristis nougareti ARAMBOURG, 1940 ‑ Maastr.: Wadi-Um-Ghudar, Jordan; Ouled Abdoun, Morocco; Iraq; Syria; Zaire

Dalpiazia stromeri CHECCHIA-RISPOLI, 1933 ‑ Maastr.: Khouribga, Morocco; Libya; Niger; Charquieh, Syria; Zaire

Ganopristis leptodon ARAMBOURG, 1935 - Maastr.: Khouribga, Morocco; Cantabria, Spain; Maastricht; Neth.; Belg.; Jordan;

Kneifiss, Syria

Ischyrhiza avonicola ESTES, 1964 ‑ Tur.: Socorro Co., NM; Tur./Cen.: Navajo Co., AZ; Con.: Ellis Co., TX;

Camp.: Newcastle Co.,DE ;Blaine Co., MT; Brewster/Delta/Lamar Co., TX; Maastr.: Weld Co., CO; McCone Co., MT;

Hunt Co., TX; Niobrara Co., WY; NC; ND; NJ; SD; Belg.; Iraq; Tunisia; Mid.Cret.: CO; Pal.: McCone Co., MT

basinensis CASE, 1987 - Up.Camp.: Washakie Co., WY

hartenbergeri CAPPETTA, 1975 - Maastr.: Pajcha Pata/Toro-Toro, Bolivia

mira (LEIDY, 1856) - Tur.: Socorro Co., NM; Grant Co., SD; San.: Chattahoochee Co., GA; Bernalillo Co., NM;

Camp.: Wilcox Co., AL; Newcastle Co.,DE ; Blaine Co., MT; Sandoval Co., NM; Fanin Co., TX; Monmouth Co., NJ;

Big Horn River, WY; AK; Alberta, Canada; Rincon Colorado, Mexico; Maastr.: Hunt Co., TX; ND; Tha.: NJ;

Low.Eoc.: Rio Arriba Co., NM; Up.Eoc.: Clarke Co., AL; Nuevo Leon, Mexico

monasterica CASE & CAPPETTA, 1997 - Maastr.: Hunt Co., TX

nigeriensis (TABASTE, 1963) - Maastr.: Mont Igdaman, Niger

schneideri SLAUGHTER & SCHNEIDER, 1968 – Cen./Tur.: Navajo Co., AZ; Tur./San.: Sandoval Co., NM; TX

ssp. – Camp.: Blaine Co., MT; Kane Co., UT; Tha./Ypr.: Dormaal, Belg.; Lut.: Natchitoches Pa., LA

viaudi CAPPETTA, 1980 – Low. San.: Vendee, France

Kiestus "Ischyrhiza" texana CASE & CAPPETTA, 1999 - Tur.: Hunt Co., TX; Maastr.: ND

Libanopristis hiram (HAY, 1903) - Cen.: Hakel, Lebanon; Alb.: Goulmirna, Morocco

Marckgrafia libyca WELLER, 1935 ‑ Cen.: Baharije, Egypt; Garaa es Sabaa, Morocco

Micropristis germaniae (WEILER, 1964) ‑ San.: Lonzee, Belg.; Camp.: Westfalen, Germany

solomonis (HAY, 1903) - Cen.: Hadjula, Lebanon

Onchopristis dunklei MCNULTY & SLAUGHTER, 1972 ‑ Low.Brm.: Teruel, Spain; Cen./Alb: Tarrant Co., TX;

Cen./Tur.: Navajo Co., AZ; Cen.: Itasca Co., MN; Lussant, France; Camp.: Brewster Co., TX; Up.Cret.: Keewatin, MN

numidus (HAUG, 1905) ‑ Cen.: Bahariya, Egypt; Draa Ubari, Libya; Taouz, Morocco; Tunisia; Alb.: Algeria

praecursor THURMOND, 1971- Aptian/Alb.: TX

ssp. - Low.Brm.: Teruel, Spain; Up.Cret.: India; N.Z.

Onchosaurus pharao (DAMES, 1887) - Tur.: Niger; San.: Gizah, Egypt; Angola; Brazil; Zaire; Camp.: TX

radicalis GERVAIS, 1852 - Sen.: Meudon, France; Tur.: Ecuador

marocanus - Maastr.: Khourigba, Morocco

sp. – San.: Pence, Bernalillo Co., NM

Plicatopristis strougoi CAPPETTA, 1991 - Maastr.: Egypt; Charquieh, Syria

Ptychotrygon agujaensis MCNULTY & SLAUGHTER, 1972 – Camp.:Big Bend, TX; Chihuahua, Mexico;

Up.Cret.: Brewster Co., TX

blainensis CASE, 1978 ‑ Camp.: Low.Brm.: Teruel, Spain; Blaine Co., MT; Lamar Co., TX; Alberta, Canada

boothi CASE, 1987 - Up.Camp.: Sandoval Co., NM; Washakie Co., WY

chattahoochiensis CASE, SCHWIMMER, BORODIN & LEGGETT 2001 - San.: Chattahoochee Co., GA

cuspidata - Up.Cret.: Monmouth Co., NJ

ellae CASE, 1987 – Up.Camp.: Sandoval Co., NM; Washakie Co., WY

eutawensis CASE et al, 2001 - San.: Chattahoochee Co., GA

greybullensis CASE, 1987 - Camp.: Washakie Co., WY

geyeri KRIWET, 1999 - Up.Alb.: E.Spain

henkeli WERNER, 1987 - Up.Cen.: Bahariya, Egypt

ledouxi CAPPETTA, 1975 - Tur.: SD

“Ptychotrygonoides” pouiti (LANDEMAINE, 1999) ‑ Cen.: Lussant, France

robyae WILLIAMSON, KIRKLAND & LUCAS, 1993 - Up.Cen./Mid.Tur.: Black Mesa, Navajo Co., AZ

slaughteri CASE, & CAPPETTA, 1975 ‑ Camp.: Fanin Co., TX

ssp. – Cen.: Brezina, Czeck.; Low.San.: Vendee, France; San.: Chattahoochee Co., GA; Wadi-Um-Ghudar, Jordan;

San./Camp.: Peru; Camp.: Sandoval Co., NM; Brewster Co., TX; Kane Co., UT; Maastr.: Monmouth Co., NJ; Morocco;

Up.Cret.: Pueblo, CO; Worland, WY

texana (LERICHE, 1940) - Maastr.: Hunt Co., TX

triangularis (RUESS, 1844)- Cen./Tur.: Navajo Co., AZ; Tur.: Socorro Co., NM; Grayson Co., TX; Bohemia, Czeck.;

Con.: Denton Co., TX; San.: Chattahoochee Co., GA; Con./San.: Bernalillo/Sandoval Co., NM;

Camp.: San Juan Co., NM; Maastr.: ND

vermiculata CAPPETTA 1975‑ Camp.: Newcastle Co.,DE ; Stewart Co., GA; Wayne Co., NC;

Camp./Maastr.: Monmouth Co., NJ; Hunt Co., TX

winni CASE & CAPPETTA, 1997‑ Maastr.: Hunt Co., TX

Pucapristis branisi SCHAEFFER, 1963 ‑ Up.Sen.: Salta Prov., Argentina; Maastr.: Potosi/Toro Toro, Bolivia

Renpetia labiicarinata WERNER, 1989 - Up.Cen.: Bahariya, Egypt

Schizorhiza stromeri WEILER, 1930 – Up.Camp./Maastr.: Coahuila, Mexico; Peru; Maastr.: Youssoufia, Morocco; Angola;

Pajcha Pata, Bolivia; Chile; Egypt; Iraq; Libya; Niger; Nigeria; Charquieh, Syria; Zaire

weileri (SERRA, 1933)- Maastr.: TX; Italy

Sclerorhynchus atavus WOODWARD, 1889 ‑ Cen.: Sahel Alma, Lebanon

fanninensis CASE & CAPPETTA, 1999 - Camp.: Fanin Co., TX

germaniae ALBERS & WEILER, 1964 – Camp.: Aachen, Germany

leivisi - Cen.: Hakel, Lebanon

pettersi CASE & CAPPETTA, 1997 ‑ Camp.: Fanin Co., TX; Maastr.: Monmouth Co., NJ; Hunt Co., TX;

Tha./Ypr.: Dormaal, Belgium

priscus CASE & CAPPETTA, 1999 - Tur./Con.: Dallas Co., TX

ssp. - Camp.: Newcastle Co.,DE ; Monmouth Co., NJ; San Juan Co., NM; Lamar Co., TX; Westfalen, Germany;

Up.Cret.: Madagascar; Morocco

Texatrygon "Ptychotrygon" (hooveri MCNULTY & SLAUGHTER, 1972) - Tur.: Grayson Co., TX; Con.: Denton Co., TX;

Camp.: Monmouth Co., NJ; Fanin Co., TX

copei CASE & CAPPETTA, 1999 - Camp.: Fanin Co., TX

Urolophidae GRAY, 1851 [Round Stingrays]

Urolophus crassicouda BLAINEVILLE, 1818 - Ypr.: Monte Bolca, Italy

cruciatus LACEPEDE, 1804 – Up.Eoc.: Wilkinson Co., GA

halleri FITCH, 1964 - Pleist.: Orange Co., CA

Myliobatiformes COMPAGNO, 1973

Dasyastidae JORDAN, 1888 [Sting Rays]

Aegyptobatus kuchnei WERNER, 1987 - Up.Cen.: Bahariya, Egypt

Arechia armbourgi CAPPETTA. 1983- Ypr.: Morocco

priemi - Tha.: Morocco

ssp. - Dan./Ypr.: Morocco; Low.Eoc.: Senegal; Mid.Eoc.: Togo

Coupatezia ambroggi CAPPETTA & NOUBHANI, 1995 - Ypr.: Morocco

boujoi NOUBHANI & CAPPETTA, 1997 - Eoc.: Khourigba, Morocco

chariri NOUBHANI & CAPPETTA, 1997 - Dan.: Morocco

danica NOUBHANI & CAPPETTA, 1997 ‑ Dan.: Youssoufia, Morocco

elevata NOUBHANI & CAPPETTA, 1997 - Maastr.: Morocco

fallax (ARAMBOURG, 1952 )‑ Maastr.: Eben Emael, Belgium; Maastricht, Neth.; Morocco

laevis NOUBHANI & CAPPETTA, 1997 ‑ Dan.: Youssoufia, Morocco

larivei NOUBHANI & CAPPETTA, 1997 ‑ Tha.: Khouribga, Morocco

melittiana NOUBHANI & CAPPETTA, 1997 - Dan.: Morocco

reniformis NOUBHANI & CAPPETTA, 1997- Maastr.: Morocco

ssp. – Maastr.: Charquieh, Syria; Tha.: Ingall, Niger; France; Bart.: BajaCalifornia, Mexico

turneri CASE & CAPPETTA, 1997 - Maastr.: Hunt Co., TX

woutersi CAPPETTA, 1982 ‑ Tha../Ypr.: Meridian, MS; Ypr.: Stafford Co,. VA; MS; Khouribga, Morocco; Egem, Belg.;

Lut.: Vald’Oise, France; Kizylkum Desert, Uzbekistan; Mid:Eoc.: Bracklesham, England

Dasyatis africana (DARTEVILLE & CASIER, 1959) - Pal.: Mandurim, Cabinda; Plio./Pleist.: Lake Turkana, Kenya

americanum HILDEBRANDT & SCHROEDER, 1924 – Low.Mio.: Beaufort Co., NC

borodini CASE, 1981 - Up.Eoc., Twiggs Co., GA

branisai CAPPETTA, 1975 - Maastr.: Pajcha Pata/Toro-Toro, Bolivia

cavernosa (PROBST, 1877) - Rup.: Espenhein, Germany; Low.Mio.: Craven Co., NC; Mid.Mio.: Loupian, France

centroura (MITCHELL, 1814) – Plio.: Beaufort Co., NC

charlisae CASE, 1981 - Up.Eoc., Twiggs Co., GA

commercensis CASE & CAPPETTA, 1997 ‑ Maastr.: Hunt Co., TX

concavifoveus CVANCARA & HOGANSON, 1993 – Pal.: ND

crosswickense CASE, 1996 - Mon.: Monmouth Co., NJ

datasi NOUBHANI & CAPPETTA, 1997‑ Ypr.: Khouribga, Morocco

davisi CASIER, 1966 - Ypr.: Sheppey, England

delfortriei CAPPETTA, 1970 - Rup.: Espenhein, Germany; Mid.Mio.: Loupian, France

duponti (WINKLER, 1874) - Ypr.: Khourigba, Morocco; Bart.: Ronquerolles, France

hexagonalis ARAMBOURG, 1952 – Mon./Dan.: Youssoufia, Morocco

jaekeli (LERICHE, 1905) ‑ Tha../Ypr.: Meridian, MS; Ypr.: Egem, Belg.; Mid:Eoc.: Bracklesham, England;

Lut.: Vald’Oise, France

martini NOUBHANI & CAPPETTA, 1997- Maastr.: Morocco

minuta CAPPETTA, 1970 - Mio.: Montpellier, France

molinoensis CAPPETTA, 1975 - Maastr.: Toro-Toro, Bolivia

muracatit (VOLTA, 1796) - Low.Eoc.: Italy

newegyptensis CASE, BORODIN & LEGETT, 2001 - Up.Maastr.: Monmouth Co., NJ

nipponensis HATAI & KATAKA, 1962 - Low.Mio.: Gifu, Japan

ponsi NOUBHANI & CAPPETTA, 1997 ‑ Pal..: Khouribga, Morocco

probsti CAPPETTA, 1970 - Mio.: Montpellier, France

puercensis WILLIAMSON & LUCAS, 1993 - Pal.: Willow Wash, San Juan Co., NM; Garfield Co., MT

“Heliobatis” radians (MARSH, 1877) – Low.Eoc.: Lincoln Co., WY

rafinesquei KUMAR & LOYAL, 1987 – Ypr.: Himachal Pradesh, Solan District, India

rugosa (PROBST, 1879) ‑ Mio.: Luble, France; Poland; Portugal; Up.Mio.: N.Germany

ruselli CAPPETTA, 1972 - Tha.: Sesso, Niger

say (LESUER, 1817) – Low.Mio.: Beaufort Co., NC

schaefferi CAPPETTA, 1975 - Maastr.: Pajcha Pata/Toro-Toro, Bolivia

serralherroi CAPPETTA, 1970 - Mio.: Montpellier, France; Portugal

sessaoensis CAPPETTA, 1972 - Tha.: Sesso, Niger

ssp. ‑ Cen.: TX; Maastr.: Carbon Co., WY; ND; Nafud, Saudia Arabia; Bolivia, Peru; Tha../Ypr.: Morocco; Pal.: San Juan Co.,

NM; MT; ND; Carbon Co., WY; Ingall, Niger; Eoc.: WY; Lut.: Covington Co., GA; Kizylkum Dsrt., Uzbekistan; Mid.Eoc.: Gerecse,

Hungary; Eoc./Plio.: BajaCalifornia, Mexico; Olig.: Gellik, Belg.; N.Z.; Low Mio.: Kent Co., DL; Effingham Co., GA;

Mio.: Kern Co., CA; Alachua Co., FL; Montpellier, France; Mizunami, Japan; Sabinata, Panama; Alicante, Spain;

Up.Mio.: Liessel, Neth.; Chancellor Point, VA; Entre Rios Prov., Argentina; Up.Mio./Low.Plio.: NE Venezuela;

Plio.: San Diego Co., CA; S.Australia; Pleist.: Contra Costa Co., CA; Hillsborough Co., FL; Holocene: Aichi, Japan

speetonensis UNDERWOOD, MITCHELL & VELTKAMP, 1999 - Hau.: Speeton, Yorkshire, England

sudrei CAPPETTA, 1972 - Pal.: Ingall, Niger

tetraedra ARAMBOURG, 1952 ‑ Dan.: Youssoufia, Morocco

thieryi SMITH, 1999 - Tha./Ypr.: Dormal/Egem, Belg.

torrejonius WILLIAMSON & LUCAS, 1993 – Pal.: Mammalon Hill, San Juan Co., NM

tricuspidatus CASIER, 1946 - Tha../Ypr.: Meridian, MS; Ypr.: Egem, Belg.; Lut.: Vald’Oise, France;

Mid.Eoc.: Bracklesham, England

vicaryi – Ypr.: Solan Dist., India

wochadunensis WARD, 1979 - Ypr.: Essex, England

Dasyrhombodus bondoni (ARAMBOURG, 1952) - Maastr.: Morocco

Delpitia reticulata NOUBHANI & CAPPETTA, 1997‑ Pal..: Khouribga, Morocco

Distobatus “Protohypolophites” morlati (WERNER, 1989) ‑ Cen.: Lussant, France; Draa Ubari, Libya

nutiae WERNER, 1987 - Up.Cen.: Bahariya, Egypt; Cen.: Gara es Sabaa, Morocco

Glickmania tamdyensa CASE, 1996 - Lut.: Kizylkum Desert, Uzbekistan

Heterobatis talbaouii NOUBHANI & CAPPETTA, 1997 ‑ Tha.etian: Stafford Co., VA; Khouribga, Morocco

Hypolophodon sylvestris WHITE, 1931 ‑ Mon.: Monmouth Co., NJ; Tha.: Marne, France; Pal.: ND; Tha./Ypr.: Dormaal, Belg.;

Ypr.: Stafford Co.,VA; Pal./Eoc.: Abbey Wood, England; Lut.: Vald’Oise, France

ssp. - Tha.: Morocco; Mid.Eoc.: Pamunkey River, VA; Lut.: Kizylkum Desert, Uzbekistan

ypresiensis CASIER, 1946 - Eoc.: Egem, Belg.

Hypolophites hutchinsi CASE, 1996 - Mon.: Monmouth Co., NJ

mayumbensis LERICHE, 1913 - Pal.: Cabinda

myliobatoides STROEMER, 1910 - Pal.: Mali; Nigeria; Togo

thaleri CAPPETTA, 1972 - Tha.: Sessao, Niger

Ishaquia globidens (ARAMBOURG, 1952) – Mon./Dan.: Tanout/Youssoufia, Morocco

Merabatis praealba (ARAMBOURG, 1952) - Eoc.: Khourbga, Morocco; ?Mio.: Beaufort Co., NC

Meridiani convexa CASE, 1994 - Low.Eoc.: Stafford Co.,VA; MS

"Dasyatis" globidens (ARAMBOURG, 1952) - Ypr.: Morocco

Myliodasyatis meskalensis NOUBHANI & CAPPETTA, 1997 - Dan.: Morocco

Prosopodon assafai NOUBHANI & CAPPETTA, 1997 ‑ Dan.: Youssoufia, Morocco

Subathanura casieri KUMAR & LOYAL, 1987 – Ypr.: Himachal Pradesh, Solan District, India

Texabatis corrugata CASE & CAPPETTA, 1997 ‑ Maastr.: Hunt Co., TX

Trygon sinhaleyos DERANIYAGAL, 1969 – Mio.: Kern Co., CA; Kala River, Sri Lanka

Viperucullus kuechnei CASE, 1996 - Mon.: Monmouth Co, NJ

Mobulidae GILL, 1893 [Devil or Manta Rays]

Archaeomanta melenhorsti HERMAN, 1979 ‑ Ypr.: Meridian, MS; Stafford Co., VA; Khouribga, Morocco; Lee‑on‑the‑Solent,

England; Egem, Belg.; Lut.: Kizylkum Desert, Uzbekistan

priemi HERMAN, 1979 - Pal./Ypr.: Morocco

ssp. ‑ Dan.: Youssoufia, Morocco; Eoc.: Guineau-Bissau; Togo

Burnhamia daviesi (WOODWARD, 1889 ‑ Tha.: Morocco; Ypr.: Stafford Co., VA; Meridian, MS;

Egem, Belg.; Burnham, England; Lut.: Vald’Oise, France Kizylkum Desert, Uzbekistan

fetahi CAPPETTA, 1979 ‑ Ypr.: Meridian, MS; Stafford Co., VA; Khouribga, Morocco

ssp. – Tha./Ypr.: Dormaal, Belg.; Ypr.: Morocco; Lut.: Covington Co., AL; Vald’Oise, France; Kizylkum Desert, Uzbekistan;

Aqt.: BajaCalifornia, Mexico

woodwardi ‑ Eoc.: Mangyshlak, Kazakhstan

Cretomanta canadensis CASE, TOKYARK & BAIRD, 1990 - Cen.: Baca Co., CO; Cen./Tur.: Russel Co., KS; Butte, Fall River Co.,

SD; Tur./Con.: TX; Con.: Saskatchewan, Canada; Maastr.: Morocco

Eomobula stehmanni HERMAN et al, 1988 - Ypr.: Stafford Co., VA; Egem, Belg.

Eoplinthicus yazooensis CAPPETTA & STRINGER, 2002 – Up.Eoc.: Copenhagen, Caldwell Parish, LA

Mobula “Manta” “Paramobula” fragilis (CAPPETTA, 1970) - Mio.: Beaufort Co., NC; Loupian, France

hynei BOURDON, 1999 - Mio.: Beaufort Co., NC

loupianensis (CAPPETTA, 1970) - Mid.Mio.: Loupian, France; Mio.: Beaufort Co., NC

melanye CASE, 1980 - Low.Mio.: Craven Co., NC

pectinata CAPPETTA, 1970 - Mio.: Beaufort Co., NC; Montpellier, France

ssp. - Olig./Mio.: Bottoms Bridge, VA; Low.Mio.: Paraguana, Venezuela; Mio.: Bakersfield, CA; Beaufort Co., NC;

Montpellier, France; Mizunami,Japan; BajaCalifornia, Mexico

Plinthicus stenodon COPE, 1869 ‑ Olig.: Berkeley Co., SC; Low.Mio.: Effingham Co., GA; NW Venezuela; Mio.: Beaufort Co., NC;

Cumberland Co., NJ; Bottoms Bridge, VA; Montpellier, France

kruibekensis BOR, 1990 ‑ Rup.: Espenhein, Germany; Mio.: USA; Up.Mio.: Liessel, Neth.; N.Germany

sp. - Mio.: FL

Myliobatidae BONAPARTE, 1838 [Eagle Rays]

Aetobatus arcuatus AGASSIZ, 1843 ‑ Low.Mio.: Domo de Zaza, Cuba; Mio: Charles Co., MD; Channey, France; Okayama, Japan;

Sabinata, Panama; Alicante, Spain; Belg.; Switzerland; Mio./Low.Plio.: Cubagua, Venezuela

irregularis (AGASSIZ, 1843) ‑ Tha../Ypr.: Morocco; Ypr.: Stafford Co,VA; MD; Sheppey, England; Lut.: Balegem, Belg.;

Guitry, France; Kizylkum Desert, Uzbekistan; Bart.: Ronquerolles, France;

Mid:Eoc.: Bracklesham, England; Birket-el-Quron, Egypt; Gerecse, Hungary

narinari (EUPHRASEN, 1790) - Pleist.: Hillsborough Co., FL; Up.Pleist.: Chatham Co., GA

rectis – Mio.: England

smithii – Mio.: Kern Co., CA

ssp. - Eoc.: GA; FL; Birket-el-Quron, Egypt; Toru Rosu, Romania; Lut.: Covington Co., AL; Mid:Eoc.: Duplin Co., NC;

Bracklesham, England; Mid.Eoc./Plio.: BajaCalifornia, Mexico; Low.Mio.: Kent Co., DL; Mio.: Craven Co., NC; FL,

Montpellier, France; Ecuador; England

sulcatus AGASSIZ, 1843 - Eoc.: Egem, Belg.

Aetomylaeus sp. (GARMAN, 1908) - Ypr.: Stafford Co., VA; Mio.: Beaufort Co., NC; Alachua Co., FL; Plio.: Montpellier, France

Aktaua kizylkumensis CASE, et al 1996 - Lut.: Kizylkum Desert, Uzbekistan

Brachyrhizodus wichitaensis ROMER, 1942 – Camp.: St. Geoges, DL; Wayne Co., NC; Camp./Maastr.: Monmouth Co., NJ;

Maastr.: Hunt Co., TX; Berivotra, NW Madagascar

Enantiobatis tarrantensis CASE & CAPPETTA, 1999 - Cen.: TX

Garabatis atlasi CAPPETTA, 1993 - Tha.etian: Ovarzazte, Morocco

Igdabatis indicus PRASSAD & CAPPETTA, 1993 - Maastr.: Asifabad, India; Lleida, Spain

sigmodon CAPPETTA, 1972 ‑ Maastr.: Kao, Niger

Ixobatis mucronata (ARAMBOURG, 1952) ‑ Maastr.: Youssoufia,, Morocco

Leidybatus granulosus CAPPETTA, 1986 ‑ Eoc.: Khouribga, Morocco

jugosus LEIDY, 1877 ‑ Ypr.: Egem, Belg. ; Lut.: Vald’Oise, France

sp. - Lut.: Vald’Oise, France

Lophobatis phosphaticus CAPPETTA, 1986 - Ypr.: Morocco

sp. - Mid:Eoc.: Bracklesham, England

Myliobatis californica GILL, 1865 – Pleist.: Orange/San Diego Co., CA

copeanus - Pal..: Charles Co., MD; Eoc.: Belvidere Beach, VA

dixoni AGASSIZ, 1843 – Pal./Eoc.: Cabinda; Tha.: France; England; Chesapeake Bay; Tha../Ypr.: Meridian, MS; Dormaal,

Belg.; Ypr.: Stafford Co., VA; Eoc.: Khouribga, Morocco; Belg.; Mid:Eoc.: Bracklesham, England; Lut.: Guitry, France

gigas COPE, 1867 - Mio.: Charles Co., MD

frangens CLARKE, 1904 - Mio.: Charles Co., MD

jugosus LEIDY, 1877 – Tha.: NJ; Mio.: Belg.

latidens WOODWARD, 1888 - Ypr.: Stafford Co., VA; Sheppey, England; Bart.: Ronquerolles, France;

Mid.Eoc.: Bracklesham, England magister COOK & RAMSDELL, 1991 – Tha.: NJ

meridionalis – Mio.: Seville, Spain; France merriami – Mio.: Kern Co., CA

monnieri CAPPETTA, 1986 ‑ Ypr.: Khouribga, Morocco

nzadinensis DARTEVELLE & CASIER, 1943 - Pal.: Cabinda, W.Africa; Ypr.: Stafford Co., VA

obesus COOK & RAMSDELL, 1991 – Tha.: NJ

oligocaena LERICHE, 1910 - Olig.: Belg.

pachydon COPE, 1867 - Mio.: Charles Co., MD

raouxi ARAMBOURG, 1953 ‑ Pal../Ypr.: Khouribga, Morocco

rivieri SAUVAGE, 1880 - Brt.: France

serratus MEYER, 1843 - Olig.: Espenheim, Germany

sinhaleyus DERANIYAGALA, 1969 - Mio.: Kala River, Sri Lanka

ssp. ‑ ?Camp.: TX; Maastr.: Nafud, Saudia Arabia; Dan.: Morocco; Dan./Plio.: BajaCalifornia, Mexico; Tha.: Meridian, MS;

Berkeley Co., SC; NJ; Pal.: ND; Germany; Tha../Ypr.: Oise, France; Ypr.: Sheppey, England; Stafford Co., VA; India;

Lut.: Covington Co., AL; Natchitoches Pa., LA; Kizylkum Dsrt,Uzbekistan; Eoc.: Twiggs Co., GA; FL; Seymour Island;

Antartica; Fayum, Egypt; Tornu Rosu, Romania; Belg.; France; Russia; Mid:Eoc.: Duplin Co., NC; Bracklesham,

England; Gerecse, Hungary; S.Australia; Prb.: Kurgan, Kazakhstan; Olig.: Isle of Wight, England; Essone, France;

Mio./Plio.: Victoria, Australia; Cape Province, South Africa; Argentina; Mio.: Alachua Co., FL; Craven Co., NC; Montpellier,

France; Malta; Gudar, Panama; Cuba; Spain; Uruguay; Up.Mio.: Liessel, Neth.; N. Germany; Mio./Low.Plio.: Cubagua, Venezuela;

Plio.: San Diego Co., CA; Peru; Pleist.: Hillsborough Co., FL

striatus BUCKLAND, 1837 - Ypr.: Stafford Co., VA; Moulares, Morocco; Bart.: France; Lut.: Vald’Oise, France;

Mid.Eoc.: Bracklesham, England; Eoc.: Balegem, Belg.

sulcidens DARTEVELLE & CASIER, 1943‑ Tha.etian/Ypr.: Khouribga, Morocco; France

toliapicus AGASSIZ, 1843 – Ypr.: Sheppey, England; Mid.Eoc.: Bracklesham, England; Lut.: Vald’Oise, France;

Mio.: Moulares, Spain

Pseudoaetobatus casieri CAPPETTA, 1986 – Lut.: Vald’Oise, France; Eoc.: Khouribga, Morocco

sp. - Ypr. Stafford Co., VA

Pteromylaeus bovinas (ST. HILLAIRE, 1817)- Mio.: Beaufort Co., NC; Montpellier, France; Low.Plio.: Cubagua, Venezuela

Zygzabatis mariae MENDIOLA CASTO, 1999 - ?Ypr.: Bou Jniba, Ouled Abdoun, Morocco

maroccana MENDIOLA CASTO, 1999 - ?Ypr.: Bou Jniba, Ouled Abdoun, Morocco

Rhinopteridae JORDAN & EVERMANN, 1896 [Cow-Nosed Rays]

Apocopodon sericeus COPE, 1886 - Pal.: Pernambuco, Brazil

Rhinoptera bonasus (MITCHELL, 1815) - Olig./Mio.: NC; Pleist.: Hillsborough Co., FL

aff. brasiliensis (MUELLER, 1936) - Olig./Mio.: NC

daviesi WOODWARD, 1889 – Ypr.: Ouled Abdoun, Morocco; Up.Eoc.: Twiggs Co., GA; Low.Mio.: Craven Co., NC

prisca WOODWARD, 1907- Pal.: Pernambuco, Brazil

raeburni WHITE, 1934 ‑ Dan.: Youssoufia, Morocco; Tha.etian: Nigeria

sherboni WHITE, 1926 ‑ Tha../Lut.: Morocco; Eoc.: Lee‑on‑the‑Solent, England;

Ypr.: Stafford Co., VA; Lut.: Vald’Oise, France; Kizylkum Dsrt., Uzbekistan;

ssp. - Up.Cret.: Holmdel, NJ; Tha.: Meridian, MS; Berkeley Co., SC; NJ; Ypr.: Morocco; Eoc.: AL; Olig.: Espenheim, Germany;

Low Mio.: Kent Co., DL; Mio.: FL; Ishikawa, Japan; Alicante, Spain; Mio./Low.Plio.: Cubagua, Venezuela;

Cape Province, South Africa;Plio.: Beaufort Co., NC; Kallo, Belg.

studeri (AGASSIZ, 1843) - Low.Mio.: Loupian, France; Switzerland

Rhombodontidae CAPPETTA, 1987

Pucabatis hoffstetleri CAPPETTA, 1975 ‑ Maastr.: Pajcha Pata/Toro-Toro, Bolivia

Rhombodus andriesi CAPPETTA & NOUBHANI, 1994 ‑ Maastr: Foum Tizi, Morocco

binkhorsti DAMES, 1881 ‑ Maastr: Monmouth Co., NJ; South Sulphur River, TX; Ruseita, Jordan; Khouribga, Morocco;

Trevino, Spain; Brazil; Chile; Jordan; Neth.; Zaire; Cret.: Wayne Co., NC

bondoni ARAMBOURG, 1952 - Maastr.: Ouled Abdoun, Morocco

kirklandi – Eocene: WY

laevis CAPPETTA & CASE, 197 5‑ Maastr.: Bowie, MD; Wayne Co., NC; Monmouth Co., NJ

meridionalis ARAMBOURG, 1952 ‑ Maastr.: Oued Erguita, Morocco; Syria

microdon ARAMBOURG, 1952 ‑ Maastr.: Youssoufia, Morocco

ssp. ‑ Camp.: Newcastle Co.,DE ; Maastr.: Hunt Co., TX; Asifabad, India; El Rosario, Mexico; Nafud, Saudia Arabia;

Lleida, Spain; Charquieh, Syria

Torpediniformes BUEN, 1926 [Electric Rays]

Narcinidae GILL, 1862

Narcine molini JAEKEL, 1894 - Ypr.: Monte Bolca, Italy

sp. - Ypr.: Morocco; Senegal; Mid.Eoc.: Togo

Pteroplatea (?Gymnura) sp.- Mio.: Montpellier, France

Torpedinidae BONAPARTE, 1838

Eotorpedo hilgendorfi (JAEKEL, 1904) - Tha../Ypr.: Khouribga, Morocco; Cameroon; Niger; Tunisia

jackeli (WHITE, 1934) - Tha../Ypr.: Meridian, MS; Morocco; Up.Pal.: Gada, Nigeria

nolfi HERMAN, 1975 - Eoc.: Stafford Co., VA; Belg.

zennaroi CAPPETTA, 1988 - Dan.: Morocco

Heterotorpedo brahimi CAPPETTA & NOUBHANI 1997‑ Tha.: Khouribga, Morocco

fowleri WARD, 1983 ‑ Ypr.: Stafford Co., VA; Mid.Eoc.: Bracklesham, England; Eoc.: France; Morocco

lyazidii CAPPETTA & NOUBHANI, 1997‑ Pal..: Khouribga, Morocco

ssp. ‑ Tha.: Ressons, France; Morocco; Tahout, Niger; Eoc.: Stafford Co., VA; Tunisia

Torpedo “Raja” dormaalensis SMITH, 1999 - Tha./Ypr.: Dormaal, Belgium

torpedo (LINNAEUS, 1758) - Plio.: S.France

sp.: Mio.: Bonpas, France